2024年2月6日發(fā)(作者:任宏偉)
廣西報春苣苔屬一新種--泡葉報春苣苔
陸仕念;符龍飛;梁桂友;溫放
【摘 要】A new species ,Primulina bullata S .N .Lu & Fang Wen ,which
grows in limestone hills and limestone caves from Guangxi is described
and illustrated .It is similar to P .liboensis(W .T .Wang &
D .Y .Chen)Mich .M?ller & A .We‐ber and
P .gueilinensis(W .T .Wang)Y .Z .Wang ,but iffers above‐mentioned two
relatives by having erect pubescence on both leaf blade surfaces ,upper
surface of leaf blade with distinctly rounded ,bullate projections ,(1)2 -9 or
more cymes , longer peduncle ,(4 .0)8 .5 - 15 cm ,linear or linear‐lanceolate
bracts ,ca .7 mm × 1 mm ,largerflower ,4 .2 -5 .3 cm long , purplish‐pink
corolla ,staminodes longer ,13 -15 mm long ,flowering time in November .%
報道了分布于廣西石灰?guī)r山地及巖溶洞穴的苦苣苔科 Gesneriaceae 報春苣苔屬
Primulina Hance 一新種———泡葉報春苣苔 P .bullata S .N .Lu & Fang Wen .泡葉報春苣苔與荔波報春苣苔 P .liboensis(W .T .Wang& D .Y .Chen)Mich .M迸ller & A .Weber 和桂林報春苣苔 P .gueilinensis(W .T .Wang)Y .Z .Wang 相近,以葉兩面均具直立柔毛,葉上面呈顯著泡狀,具較多花序,(1)2~9條,花序梗較長,(4.0)8.5~15 cm ,苞片線形或線狀披針形,大小為7 mm ×1 mm ,花大,長4.2~5.3 cm ,粉紫色,退化雄蕊較長,13~15 mm ,花期在11月而區(qū)別.
【期刊名稱】《廣西植物》
【年(卷),期】2013(000)001
【總頁數(shù)】5頁(P42-45,81)
【關鍵詞】報春苣苔屬;廣西;苦苣苔科;泡葉報春苣苔;新分類群
【作 者】陸仕念;符龍飛;梁桂友;溫放
【作者單位】靖西縣林業(yè)局 林業(yè)種苗試驗站,廣西 百色 533800;廣西師范大學,廣西 桂林 541002; 廣西壯族自治 區(qū)中國科學院廣西植物研究所,廣西 桂林
541006;廣西壯族自治 區(qū)中國科學院廣西植物研究所,廣西 桂林 541006;廣西壯族自治 區(qū)中國科學院廣西植物研究所,廣西 桂林 541006
【正文語種】中 文
【中圖分類】Q949
原唇柱苣苔屬Chirita Buch.- 是一個以我國華南、西南為分布中心的苦苣苔科Gesneriaceae大屬,我國分布有3個組,即唇柱苣苔組accus,麻葉唇柱苣苔組a和鉤序唇柱苣苔組hirita(王文采,1990;王文采等,1998;李振宇等,2004;韋毅剛等,2010)。然而在2011年對該屬的修訂中,原唇柱苣苔組以及文采苣苔屬Wentsaiboea & (僅其中的2個種文采苣苔lia
& 和羅城文采苣苔ngensis Yan Liu & )(Liu et al.,2010)以及唇柱苣苔組的近緣類群——小花苣苔屬Chiritopsis 被并入我國的一個特有單型屬——報春苣苔屬Primulina Hance內(nèi),鉤序唇柱苣苔組被提升為鉤序苣苔屬Microchirita(),而原麻葉唇柱苣苔組中大部分的種以及另一個單型屬,密序苣苔屬Hemiboeopsis 被并入漢克麗亞花屬漢克麗亞花組Henckelia lia中,少數(shù)種則并入一個重新恢復的屬——Damrongia Kerr ex Craib之中,而原唇柱苣苔屬中的ia也被提升為屬,即Liebigia Endl.(Wang et al.,2011;Weber et
al.,2011)。
廣西是原唇柱苣苔屬唇柱苣苔組,如今的報春苣苔屬的主要分布中心,但該屬除了少數(shù)種,如螞蝗七Primulina fimbripala(Hand.-Mazz.)、牛耳朵a(Hance)、神農(nóng)架報春苣苔uba()等之外,絕大部分為狹域分布的物種,常僅見于一兩個石灰?guī)r山地山頭或巖溶洞穴。作者近幾年關注于廣西石灰?guī)r山地以及石灰?guī)r巖溶洞穴的苦苣苔科植物的研究,在對這一特殊的,分布于極為狹窄的區(qū)域的該類群植物進行詳細的野外考察過程中,在廣西境內(nèi)的石灰?guī)r山地林下、巖溶洞穴分別采集到許多該科植物的標本,其中部分當時被鑒定為未詳知種。在對這些植物進行充分和長時間的野外定點觀察和生物學性狀調查,我們發(fā)現(xiàn)其中分布于靖西縣(百色市)一個報春苣苔屬物種在相關的苦苣苔科植物文獻與專著中(王文采,1990;Wang et al.,1998;李振宇等,2004;韋毅剛等,2010;Liu et al.,2010,2011;Xu et al.,2010;Tang & Wen,2011;Weber et al.,2011;Wu et
al.,2011;Xu et al.,2010,2011)均無記載,是未被描述的新分類群。
泡葉報春苣苔 新種 圖1
Primulina bullata & Fang Wen,.1 & 2.
It differs from Primulina liboensis( & )Mich.M?ller &
and nensis() (Table 1) having erect
pubescence on both leaf blade surfaces,upper surface of leaf blade with
distinctly rounded,bullate projections,(1)2-9 or more cymes,longer
peduncle,(4.0)8.5-15 cm,linear or linear-lanceolate bracts,ca.7 mm×1
mm,larger flower,4.2-5.3 cm long,purplish-pink,staminodes longer,13-15 mm long,late flowering time in Nov.
Perennial herbs. Rhizome subterete,1.5-3.5 cm long or longer,1.0-1.5
cm in diameter. Leaves 6-12 or more,all basal,three leaves verticillate;petiole compresd,cross ction mi-elliptic,2-7.5 cm long,0.6-1.1
cm wide,short puce strigo on both surfaces;blades dark-green,coriaceous or stiffly chartaceous,obliquely ovate,elliptic to broadly
oblong-ovate,left-right asymmetric or symmetric,(5)11-13.5 cm×(3.5)7-9.5 cm,upper surface distinctly bullate,cuneate at ba,commonly
symmetric,occasionally oblique,margin with 15-25 crenation on each
side,acuminate,obtu or round at apex,with erect pubescence on
both surfaces,margins with tricholoma,pubescence 0.5-1 mm long,all
veins commonly silvery white,lateral veins 4-5 on each side,impresd
adaxially and prominent abaxially. Cymes axillary,(1)2-9 or more,1-2-branched,(1)3-9 or more flowers on one cyme;peduncle(4.0)8.5-15 cm
long,slender,(1.5)1.9-2.2 mm in diameter,erect white-pubescent;bracts 2,opposite,purplish green,linear or linear-lanceolate,ca.7×1
mm,margin entire,obtu at apex,appresd purple-pubescent outside,glabrous inside. Bracteoles 2,opposite,shape,hairs and colour the
same as bracts but apex acute. Pedicel 6-11 mm long,calyx 5-parted near
to the ba,lobe narrowly lanceolate-linear,8-11×2-2.5 mm,margin
entire,acute at apex,appresd white pubescent outside,glabrous
inside. Corolla 4.2-5.3 cm long,nearly glabrous outside,but sparly
short purplish red strigo along the back of corolla,glabrous inside;tube funnel-formed,purplish-pink,2.5-3.0 cm long,0.8-1.2 cm in
diameter in medium,1.2-1.5 cm in diameter at the orifice;throat with
two yellow lines;limb distinctly 2-lipped,purple or purplish pink,adaxial
lip 2-lobed to or over the middle,lobes ovate or nearly orbicular,0.9-1.1
cm×ca.0.9 cm,obtu at apex,with two deep purple lines inside;abaxial
lip 3-lobed over the middle, two lateral lobes inapparently obliquely
ovate or oval,the central one oblong,12.5-15 mm×10.5-12 mm,rounded at apex. Stamens 2,adnate to 1.2-1.5 cm above the ba of the
corolla tube;filaments geniculate near the ba,1.3-1.5 cm long,glabrous;anthers subreniform,slightly contracted in the middle,3.5-4.5
mm long,glabrous. Staminodes 3, lateral ones 6.5-8.0 mm long,
glabrous,adnate to 12-15 mm above the ba of the corolla tube,middle one ca 0.5 mm long,adnate to ca.3-4 mm above the ba of the
corolla tube. Disc annular,1.0-1.5 mm high,margin repand or entire.
Pistil 2.8-2.9 cm long,ovary linear,ca.2.3 cm long,ca 1.4-2.0 mm in
diameter,glandular-pubescent;style ca.5-6 mm long,nearly glabrous;stigma obtrapeziform,ca.2 mm long,apex 2-lobed. Capsule not en.
Flowering in Nov.
China(中國),Guangxi(廣西):Jingxi County(靖西縣),Hurun Town(湖潤鎮(zhèn)),F(xiàn)rom Da’ai vil-lage to Dongba village(從達愛村到東巴村途中),on the
moist rock face near to road and at the entrance of limestone caves,rare,526 m a.s.l.,09 Oct.2010,F(xiàn)ang Wen & WFBCJT11110901(holotype:
IBK;isotype: BJFC).
本種與荔波報春苣苔Primulina liboensis( & )Mich.M?ller
& 和桂林報春苣苔nensis()在親緣關系上相接近,但葉兩面均具直立柔毛,葉上面呈顯著泡狀,具較多花序,(1)2~9條,花序梗較長,(4.0)8.5~15 cm,苞片線形或線狀披針形,大小為7 mm×1 mm,
花大,長4.2~5.3 cm,粉紫色,退化雄蕊較長,13~15 mm,花期11月,可以區(qū)別(表1)。
致謝 承蒙林文宏先生為本文繪圖,在此表示衷心的謝意!
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李振宇,王印政. 2004. 中國苦苣苔科植物[M]. 鄭州:河南科學技術出版社:171-261
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